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During the later

parts of copulation, more frequent behaviors were the following: the male

rested one paw on the ground or by the female's side, and he placed one or both

paws up on the female's back; the female walked and sniffed or scratched the

ground while mounted (Table 2).

When the male was not biting the female's neck, his head was either resting on

her back, pressed against her side, hung down near her side or held above her

back (Fig...

2). The female did not remain stationary for the entire copulation.

but walked several meters every few minutes. The female both initiated and

returned bites towards the male. biting his ears, head or forepaws. While

mounted, the female might walk, look around, sniff the ground, eat grass, drink

water, or stand still with her head up...

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